Wilfred Wollheim

Co-director, Water Systems Analysis Group
Phone: (603) 862-5022
Office: Natural Resources & the Environment, James Hall Rm 276, Durham, NH 03824
Pronouns: He/him/his
Wilfred Wollheim

In understanding the environment as an integrated system, WSAG explores the physical, chemical and biological processes that shape hydrological systems, with emphasis on the unique role of humans as agents of change. Our research and educational programs cross the boundaries of traditional scientific disciplines to foster interdisciplinary understanding of environmental change. Through field monitoring, whole ecosystem experiments, novel in situ sensor deployments, and dynamic hydrological and ecosystem modeling, we attempt to understand hydrological and biogeochemical dynamics at scales ranging from individual ecosystems, to whole river systems, to the global systems of inland waters. Built on strong collaborations that are national and international in scope, our group also serves as a teaching platform for the next generation of ecosystem and earth system scientists.

The success of the Water Systems Analysis Group reflects a rapidly emerging scientific field which pursues integrative studies of hydrology, biogeochemistry, and human-water interactions that is necessary to analyze the full dimension of anthropogenic change at local, regional, and global scales.


  • Ph.D., Earth Sciences, University of New Hampshire
  • M.S., Zoology/Physiology, University of Wyoming
  • B.S., Natural Resources, Cornell University

Courses Taught

  • NR 504: Freshwater Resources
  • NR 707/807: Environmental Modeling
  • NR 751/851: Aquatic Ecosystems
  • NR 791: Proj/Environmental Science I
  • NR 795W: Investigations
  • NR 905: Grant Writing
  • NR 993: Sem/ Science Denialism

Selected Publications

Robison, A. L., Wollheim, W. M., Turek, B., Bova, C., Snay, C., & Varner, R. K. (n.d.). Spatial and temporal heterogeneity of methane ebullition in lowland headwater streams and the impact on sampling design. Limnology and Oceanography. doi:10.1002/lno.11943

Covino, T. P., Wlostowski, A. N., Gooseff, M. N., Wollheim, W. M., & Bowden, W. B. (2021). The Seasonality of In‐Stream Nutrient Concentrations and Uptake in Arctic Headwater Streams in the Northern Foothills of Alaska's Brooks Range. Journal of Geophysical Research: Biogeosciences, 126(4). doi:10.1029/2020jg005949

Borsuk, M. E., Mavrommati, G., Samal, N. R., Zuidema, S., Wollheim, W., Rogers, S. H., . . . Gardner, K. (2019). Deliberative multiattribute valuation of ecosystem services across a range of regional land-use, socioeconomic, and climate scenarios for the upper Merrimack River watershed, New Hampshire, USA. ECOLOGY AND SOCIETY, 24(2). doi:10.5751/ES-10806-240211

Bernal, S., Lupon, A., Wollheim, W. M., Sabater, F., Poblador, S., & Marti, E. (2019). Supply, Demand, and In-Stream Retention of Dissolved Organic Carbon and Nitrate During Storms in Mediterranean Forested Headwater Streams. FRONTIERS IN ENVIRONMENTAL SCIENCE, 7. doi:10.3389/fenvs.2019.00060

Talbot, C. J., Bennett, E. M., Cassell, K., Hanes, D. M., Minor, E. C., Paerl, H., . . . Xenopoulos, M. A. (2018). The impact of flooding on aquatic ecosystem services. BIOGEOCHEMISTRY, 141(3), 439-461. doi:10.1007/s10533-018-0449-7

Deegan, L. A., Johnson, D. S., Warren, R. S., Peterson, B. J., Fleeger, J. W., Fagherazzi, S., & Wollheim, W. M. (2012). Coastal eutrophication as a driver of salt marsh loss. NATURE, 490(7420), 388-+. doi:10.1038/nature11533

Beaulieu, J. J., Tank, J. L., Hamilton, S. K., Wollheim, W. M., Jr, H. R. O., Mulholland, P. J., . . . Thomas, S. M. (2011). Nitrous oxide emission from denitrification in stream and river networks. PROCEEDINGS OF THE NATIONAL ACADEMY OF SCIENCES OF THE UNITED STATES OF AMERICA, 108(1), 214-219. doi:10.1073/pnas.1011464108

Harrison, J. A., Maranger, R. J., Alexander, R. B., Giblin, A. E., Jacinthe, P. -A., Mayorga, E., . . . Wollheim, W. M. (2009). The regional and global significance of nitrogen removal in lakes and reservoirs. BIOGEOCHEMISTRY, 93(1-2), 143-157. doi:10.1007/s10533-008-9272-x

Peterson, B. J., Wollheim, W. M., Mulholland, P. J., Webster, J. R., Meyer, J. L., Tank, J. L., . . . Morrall, D. D. (2001). Control of nitrogen export from watersheds by headwater streams. SCIENCE, 292(5514), 86-90. doi:10.1126/science.1056874

Mulholland, P. J., Tank, J. L., Sanzone, D. M., Wollheim, W. M., Peterson, B. J., Webster, J. R., & Meyer, J. L. (2000). Nitrogen cycling in a forest stream determined by a N-15 tracer addition. ECOLOGICAL MONOGRAPHS, 70(3), 471-493. doi:10.1890/0012-9615(2000)070[0471:NCIAFS]2.0.CO;2

Most Cited Publications