Ray Grizzle

Research Professor of Zoology

Ph.D. Rutgers University, 1988

Research Expertise

Oyster ecology, restoration and aquaculture, Intergrated multi-trophic aquaculture, benthic ecology, bivalve mollusks, seafloor habitat mapping, including remote sensing and underwater viodeography

Research Emphasis

Much of my recent research has involved restoration of oyster reefs and oyster aquaculture in New England, including integrated farming methods for oysters and seaweeds. My laboratory also has participated in a variety of seafloor mapping projects focusing on towed underwater video, but also including sonar and other remote sensing techniques. Other research topics include the effects of marine protected areas on seafloor ecology and fisheries, impacts of aquaculture activities on benthic communities, and succession of fouling communities on aquaculture cages.

Learn more about Dr. Grizzle's Oyster Restoration Program


Zu Ermgassen, P.S.E., M.D. Spalding, R.E. Grizzle and R.D. Brumbaugh. 2012. Quantifying the loss of marine ecosystem service: filtration by the eastern oyster in US estuaries. Estuaries and Coasts DOI 10.1007/s12237-012-9559-y.

Zu Ermgassen, P.S.E., M.D. Spalding, B. Blake, L.D. Coen, B. Dumbauld, S. Geiger, J.H. Grabowski, R. Grizzle, M. Luckenbach, K. MacGraw, W. Rodney, J.L. Ruesink, S.P. Powers and R.D. Brumbaugh. 2012. Historical ecology with real numbers: past and present extent and biomass of an imperiled estuarine habitat. Proceedings of the Royal Society, B DOI 10.1098/rspb.2012.0313.

Grizzle, R.E., L.G. Ward, L.A. Mayer, M.A. Malik, A.B. Cooper, H.A. Abeels, J.K. Greene, M.A. Brodeur and A.A. Rosenberg. 2009. Effects of a large fishing closure on benthic communities in the western Gulf of Maine, USA: recovery from the effects of gillnets and otter trawls. Fishery Bulletin 107:308-317.

Grizzle, R.E., J.K. Greene and L.D. Coen. 2008. Seston removal by natural and constructed intertidal eastern oyster (Crassostrea virginica) reefs: a comparison with previous laboratory studies, and the value of in situ methods. Estuaries and Coasts 31:1208-12220.

Capone, M., R. Grizzle, A.C. Mathieson and J. Odell. 2008. Intertidal oysters in northern New England. Northeastern Naturalist 15:209-214.

McDermott, S., D. Burdick, R. Grizzle and J. Greene. 2008. Restoring ecological functions and increasing community awareness of an urban tidal pond using blue mussels. Ecological Restoration 26:254-262.

Grizzle, R.E., M.A. Brodeur, H.A. Abeels and J.K. Greene. 2007. Bottom habitat mapping using towed underwater videography: subtidal oyster reefs as an example application. Journal of Coastal Research 24:103-109.

Greene, J.K. and R.E. Grizzle. 2007. Successional development of fouling communities on open ocean aquaculture fish cages in the western Gulf of Maine, USA. Aquaculture 262:289-301.

Grizzle, R.E., J.K. Greene, M.W. Luckenbach and L.D. Coen. 2006. A new in situ method for measuring seston uptake by suspension-feeding bivalve mollusks. Journal of Shellfish Research 25:643-649.

Hofmann, E.E., J.M. Klinck, J.N. Kraeuter, E.N. Powell, R.E. Grizzle, S.C. Buckner and V.M. Bricelj. 2006. A population dynamics model of the hard clam, Mercenaria mercenaria: development of the age- and length-frequency structure of the population. Shellfish Research 25:417-444.

Grizzle, R.G., L.G. Ward, J.R. Adams, S.J. Dijkstra and B. Smith. 2005. Mapping and characterizing subtidal oyster reefs using acoustic techniques, underwater videography and quadrat counts. In: P.W. Barnes and J.P. Thomas (eds.), Benthic Habitats and the Effects of Fishing. Bethesda, MD, pp. 153-159. American Fisheries Society Symposium 41.

Cutter, G.R., Jr., Y. Rzhanov, L.A. Mayer and R.E. Grizzle. 2005. Ground-truthing benthic habitat characteristics using video mosaic images. American Fisheries Society Symposium 41:171-177.

Wall, L.M., L.J. Walters, R.E. Grizzle and P.E. Sacks. 2005. Recreational boating activity and its impact on the recruitment and survival of the oyster Crassostrea virginica on intertidal reefs in Mosquito Lagoon, FL. J. Shellfish Research 24:965-973.

Grizzle, R.E., L.G. Ward, R. Langan, G.M. Schnaittacher, J.A. Dijkstra and J.R. Adams. 2003. Environmental monitoring at an open ocean aquaculture site in the Gulf of Maine: Results for 1997-2000. In: Bridger, C.J. and B.A. Costa-Pierce (eds.). Open Ocean Aquaculture: From Research to Commercial Reality. The World Aquaculture Society, Baton Rouge, Louisiana. pp. 105-117.

Grizzle, R.E., J.R. Adams and L.J. Walters. 2002. Historical changes in intertidal oyster (Crassostrea virginica) reefs in a Florida lagoon potentially related to boating activities. J. Shellfish Research 21:749-756.

Raz-Guzman, A. and R.E. Grizzle. 2001. Techniques for quantitative sampling of infauna and small epifauna in seagrass. In: Short, F.T. and R.G. Coles (eds.). Global Seagrass Research Methods. Elsevier Science B.V. Amsterdam. pp. 237-255.

Grizzle, R.E., M. Bricelj, S. Shumway. 2001. Physiological ecology of Mercenaria mercenaria. In: Kraeuter, J.N. and M. Castagna (eds.). Biology of the Hard Clam. Elsevier Sci. Publ., Amsterdam. pp. 305-382.

Grizzle, R.E., F.T. Short, C.R. Newell, H. Hoven, L. Kindblom. 1996. Hydrodynamically induced synchronous waving of seagrasses: 'monami' and its possible effects on larval mussel settlement. Journal of Experimental Marine Biology and Ecology 206:165-177.

Grizzle, R.E., R. Langan and W.H. Howell. 1992. Growth responses of suspension-feeding bivalve molluscs to changes in water flow: differences between siphonate and non-siphonate taxa. Journal of Experimental Marine Biology and Ecology 162:213-228.

Grizzle, R.E. and C.A. Penniman. 1991. Effects of organic enrichment on estuarine macrofaunal benthos: a comparison of sediment profile imaging and traditional data. Marine Ecology Progress Series. 74:249-262.


Jackson Estuarine Laboratory
85 Adams Point Road
Durham, NH 03824

Research Areas: 
Benthic Ecology
Coastal Restoration
Marine Biology
Ocean and Seafloor Mapping